The other three lineages illustrate types of conflict between molecular species delimitation and traditional morphology- based taxonomy.

A Picture Is Worth a Thousand Controversies Flame wars aside, opponents of photo- based taxonomy aren't so much worried about a fly in South Africa.

The pair not only named a new species without a specimen, but wrote that photography- based taxonomy was increasingly important in this age of high-res imaging.

Anti-photo taxonomists penned a formal response to the Pape correspondence, "Photography- Based Taxonomy Is Inadequate, Unnecessary, and Potentially Harmful for Biological Sciences"-completewith almost 500 signatories.