Meanings of further specialization in English

Within each major lineage, further specialization occurs, resulting in distinct subsets that differ in TCR specificity, phenotype and functional attributes.

It is simply a further development of a classic style, a further specialization, the homogeneous in Spencerian phraseology having become heterogeneous.

Finally, some lens crystallin genes undergo more extensive duplication, usually followed by a further specialization of some copies for lens functions, as expected.